TTO Changes

Notes to the Reader

Abbreviations

CASGEN - California Academy of Sciences, Catalog of Fishes (CoF), Genera Database

CASSPC - California Academy of Sciencies, Catalog of Fishes (CoF), Species Database

CoF - William Eschmeyer's Catalog of Fishes

TTO - Teleost Taxonomy Ontology

Possible actions for each change

Note: Obsoleting a term means it is no longer available for use and does not appear in the main hierarchy of the TTO. Phenoscape needs to retain the name and term id so that prior annotations to the term do not break.

• Family added A new TTO term is created, which should have a dbxref referring to the publication or some CoF derived identifier.
• Family obsoleted The TTO term and id are obsoleted. Genera are moved to a new family or a higher taxon.

• Genus added A new TTO term is created, which should have a dbxref referring to the publication or a (new) CASGEN identifier
• Genus obsoleted The TTO term and id are obsoleted. Any species under this genus are changed to the new genus.
• Genus moved The existing genus has its is_a relation changed to point to the new family or higher taxon.

• Species added A new TTO term is created, which should have a dbxref referring to the publication or a (new) CASSPC identifier.
• Species changed to/added as synonym of The TTO term and id are retained; the name is changed to the new name, the old name is added as a synonym. The is_a relation for the species is updated to the new genus. The synonym should have a dbxref refering to the publication. This is the default when the containing genus changes.
• Species obsoleted/added as synonym of The term is obsoleted, the old name is added as a synonym of the new name. The CASSPC or other dbxref of the old term should be added to the synonym. This is the default when species are merged.

Changes resulting from publication Fink and Fink (1981)

Citation is: Fink, SV. and Fink, WL. 1981. Interrelations of the ostariophysan fishes (Teleostei) Zoological Journal of the Linnean Society 72:297-353.

• Abramites sp. added as species
• Albula sp. added as species
• Apareiodon sp. added as species
• Astroblepus sp. added as species
• Brycon sp. added as species
• Bunocephalus sp. added as species
• Carpiodes sp. added as species
• Catostomus sp. added as species
• Characidium sp. added as species
• Clarias sp. added as species
• Coregonus sp. added as species
• Creagrutus sp. added as species
• Ctenolucius sp. added as species
• Distichodus sp. added as species
• Eigenmannia sp. added as species
• Esox sp. added as species
• Gymnocorymbus sp. added as species
• Gyrinocheilus sp. added as species
• Hemiodus sp. added as species
• Homaloptera sp. added as species
• Ichthyborus sp. added as species
• Ictalurus sp. added as species
• Megalops sp. added as species
• Mesoborus sp. added as species
• Poecilocharax sp. added as species
• Salmo sp. added as species
• Sternopygus sp. added as species
• Vandellia sp. added as species

Change from a curator request

• Danio devario added as synonym of Devario devario

Changes resulting from a project decision to add and reorganize higher level taxa to follow Nelson(2006) and the ToLWeb website

Sources are the ToLWeb website and Nelson(2006) citation for latter source is:

• Chordata added as phylum
• Myxinomorphi added without rank
• Hyperotreti added as synonym of Myxinomorphi
• Myxini removed (not obsoleted)
• Vertebrata added without rank
• Petromyzontomorphi added without rank
• Cephalaspidomorphi removed
• Gnathostomata added without rank
• Chondrichthyes added without rank
• Chimaeriformes added as order
• Osteichthyes added without rank
• Cladistia added without rank
• Polypteriformes added as order
• Chondrostei added without rank
• Neopterygii added without rank
• Halecostomi added without rank
• Teleostei added without rank
• Osteoglossomorpha added without rank
• Hiodontiformes added as as order
• Elopomorpha added without rank
• Ostarioclupeomorphia added without rank
• Otocephala added as synonym Ostarioclupeomorphia
• Clupeomorphia added without rank
• Ostariophysi added without rank
• Anotophysi added without rank
• Otophysi added without rank
• Euteleostei added without rank
• Protacanthopterygii added without rank
• Osmeriformes removed (not obsoleted)
• Neoteleostei added without rank
• Stenopterygii added without rank
• Ateleopodomorpha added without rank
• Cyclosquamata added without rank
• Scopelomorpha added without rank
• Acantomorpha added without rank
• Lampridiformes added without rank
• Acanthopterygii added without rank
• Smegmamorpha added without rank
• Elassomatiformes added as order
• Mugiliformes added as order
• Dactylopteriformes added as order
• Coelocanthiformes added as order

Changes resulting from publication Britz and Hoffman (2006)

Citation is: Britz, R. and Hoffmann M. 2006. Ontogeny and homology of the claustra in otophysan Ostariophysi (Teleostei) Journal of Morphology 267:90-923.

• Ancistrus sp. added as species
• Clarias sp. added as species
• Puntius sp. added as species
• Devario cf. aequipinnatus added as species
• Hemiloricaria beni added as synonym of Rineloricaria beni

Changes resulting from publication Fang (2003)

• Danio aff. dangila added as species
• Chela sp. added as species
• Esomus sp. added as species

Changes resulting from publication Sidlauskas and Vari (2008)

Citation is: Sidlauskas, BL. and Vari RP. Phylogenetic relationships within the South American fish family anostomidae (Teleostei, Ostariophysi, Characiformes). Zoological Journal of the Linnean Society 154:70-210.

• Laemolyta sp. added as species
• Leporinus cf. ecuadorensis added as species
• Leporinus cf. fasciatus added as species
• Leporinus cf. friderici added as species
• Leporinus cf. moralesi added as species
• Leporinus cf. niceforoi added as species
• Leporinus sp. added as species
• Citharinus sp. added as species
• Distichodus sp. added as species
• Petulanos added as genus
• Anostomus intermedius changed to/added as synonym of Petulanos intermedius
• Anostomus plicatus changed to/added as synonym of Petulanos plicatus
• Anostomus spiloclistron changed to/added as synonym of Petulanos spiloclistron
• Hypomasticus added as genus (was subgenus of Leporinus)
• Leporinus despaxi changed to/added as synonym of Hypomasticus despaxi
• Leporinus megalepis changed to/added as synonym of Hypomasticus megalepis
• Leporinus mormyrops changed to/added as synonym of Hypomasticus mormyrops
• Leporinus pachycheilus changed to/added as synonym of Hypomasticus pachycheilus

Changes resulting from invited update (April 2008) of Mayden and Conway

Notes: This update contains many subfamily assignments. We don't currently include subfamilies in the TTO, but the update file will be retained in case this policy changes.

• Paracanthocobitis added as synonym to Acanthocobitis (Grant 2007)
• Botiidae added as family (Nalbant, T. T. 2002))
• Botia moved to family Botiidae
• Chromobotia moved to family Botiidae
• Leptobotia moved to family Botiidae
• Parabotia moved to family Botiidae
• Sinibotia moved to family Botiidae
• Syncrossus moved to family Botiidae
• Yasuhikotakia moved to family Botiidae
• Bibarba added as genus to family Cobitidae (Chen & Chen 2007)
• Bibarba bibarba added as species (Chen & Chen 2007)
• Erythroculter no change (unpublished suggestion to demote to synonym of Chandoichthys)
• Akrokolioplax no change (already in CoF CASGEN:10888)
• Henicorhynchus obsoleted/demoted to synonym of Bangana (Smith 1945)
• Henicorhynchus lobatus changed to/added as synonym of Bangana lobatus
• Henicorhynchus siamensis changed to/added as synonym of Bangana siamensis
• Henicorhynchus lineatus changed to/added as synonym of Bangana lineatus
• Henicorhynchus ornatipinnis changed to/added as synonym of Bangana ornatipinnis
• Acra changed so it is now a synonym of Bangana (was subgenus in CoF)
• Longiculter no change (uncertain subfamily change)
• Chalcalburnus obsoleted/demoted to synonym of Alburnus (as per online CoF) - member species pending TTO revision
• Psilorhynchoides obsoleted/demoted to synonym of Psilorhynchus (Conway & Kottelat 2007)
• Psilorhynchoides pseudecheneis changed to /demoted to synonym of Psilorhynchus pseudecheneis
• Psilorhynchoides arunachalensis changed to/demoted to synonym of Psilorhynchus arunachalensis
• Psilorhyncus robustus added as species (Conway & Kottelat 2007)
• Celesteichthys obsoleted/demoted to synonym of Danio (Roberts 2007)
• Celesteichthys margaritatus changed to/added as synonym of Danio margaritatus
• Bengala demoted to synonym of Megarasbora
• Bengala elanga changed to/demoted to synonym of Megarasbora elanga
• Cyprinopsis no change, problem appears in description date, not covered in TTO
• Mirolabeo no change, uncertain demotion to synonym of Bangana
• Vaillantellidae added as family (Slechtova et al. 2007)
• Vaillantella moved to family Vaillantellidae
• Caucus – appears as cypriniform in online CoF, but was not in CAS species table so never added to TTO; after consultation recommendation was to remove/not add; no change.
• Oreodaimon – appears as synonym to Pseudolabrus in online CoF, no change.
• Orthrias barbatulus zetensis no change (already demoted in CoF)
• Schistura cryptofasciata no change (already in CoF; CASSPC 68781)
• Notropis harperi changed to/demoted to synonym of Pteronotropis harperi
• Microrasbora micropthalma added as species (Jiang, Chen & Yang 2008)
• Araiocypris added as genus to Cyprinidae
• Araiocypris batodes added as species (Conway & Kottelat 2008)

Changes resulting from publication Conway (2005)

• Rasbora cf. bankanensis added as species

Changes resulting from publication de Pinna and Grande (2003)

• Osteoglossum sp. added as species
• Elops sp. added as species

Changes resulting from publication Conway et al. (2008)

• Oxygaster hypophthalmus added as synonym of Parachela hypophthalmus
• Rasboroides vaterifloris added as synonym of Rasbora vaterifloris
• Devario cf. aequipinnatus added as species

Changes resulting from publication Vari(1995)

Citation is: Vari RP. 1995. The Neotropical fish family Ctenoluciidae (Teleostei: Ostariophysi: Characiformes): supra and intrafamilial phylogenetic relationships, with a revisionary study. Smithsonian Contributions to Zoology 564: 103.

• Agoniates sp. added as species
• Copella sp. added as species
• Distichodus sp. added as species
• Eugnathichthys sp. added as species
• Hemistichodus sp. added as species
• Hoplias cf. malabaricus added as species
• Hydrocynus sp. added as species
• Lebiasina sp. added as species
• Mesoborus sp. added as species
• Pyrrhulina sp. added as species
• Salminus sp. added as species
• Hydrolycus pectoralis added as synonym of Hydrolycus scomberoides

Changes resulting from publication Cavender and Coburn (1992)

Citation is: Cavender, Ted M., and Miles M. Coburn. 1992. Phylogenetic relationships of North American cyprinids. Pp. 293-327. In: Systematics, Historical Ecology, and North American Freshwater Fishes. R.L. Mayden (Ed.). Stanford University Press. Stanford, California. 969 p.

• Spratellicypris added as synonym for genus Puntius
• Hybosis labrosa added as synonym for Cyprinella labrosa
• Hybosis monocha added as synonym for Erimonax monocha
• Hybosis zanema added as synonym for Cyprinella zanema
• Notropis ardens added as synonym for Lythrurus ardens
• Notropis caeruleus added as synonym for Cyprinella caerulea
• Notropis callistius added as synonym for Cyprinella callistia
• Notropis camurus added as synonym for Cyprinella camura
• Notropis cerasinus added as synonym for Luxilus cerasinus
• Notropis chloristius added as synonym for Cyprinella chloristia
• Notropis chrysocephalus added as synonym for Luxilus chrysocephalus
• Notropis coccogenis added as synonym for Luxilus coccogenis
• Notropis formosus added as synonym for Luxilus formosa
• Notropis galacturus added as synonym for Cyprinella galactura
• Notropis niveus added as synonym for Cyprinella nivea
• Notropis pyrrhomelas added as synonym for Cyprinella pyrrhomelas
• Notropis trichroistius added as synonym for Cyrinella trichroistia
• Notropis venustus added as synonym for Cyprinella venusta
• Notropis whipplei added as synonym for Cyprinella whipplei
• Notropis xaenurus added as synonym for Cyprinella xaenura
• Notropis zonatus added as synonym for Luxilus zonatus
• Notropis zonistius added as synonym for Luxilus zonistius
• Semotilus margarita added as synonym for Margariscus margarita
• Engraulicypris argenteus added as synonym for Rastrineobola argentea
• Moroco steindachneri added as synonym for Phoxinus steindachneri
• Osteocheilus hasseltii added as synonym for Osteochilus hasseltii
• Rhotee cotio added as synonym for Osteobrama cotio

Changes resulting from publication de Pinna (1996)

Citation is: de Pinna MCC. 1996. A phylogenetic analysis of the Asian catfish families Sisoridae, Akysidae, and Amblycipitidae, with a hypothesis on the relationships of the neotropical Aspredinidae (Teleostei, Ostariophysi). Fieldiana: Zoology (New Series) 84:1-83.

• Akysis sp. 1 added as species
• Akysis sp. 2 added as species
• Akysis sp. 3 added as species
• Erethistes sp. added as species
• Exostoma sp. added as species
• Glyptothorax sp. added as species
• Bunocephalus sp. added as species
• Ernstichthys sp. added as species

Changes resulting from publication Conway and Mayden (2007)

Citation is: Conway, KW. and Mayden RL. 2007. The gill arches of Psilorhynchus (Ostariophysi: Psilorhynchidae). Copeia ? 267-280.

• Acanthocobitis sp. added as species
• Beaufortia cf. leveretti added as species
• Gastromyzon sp. added as species
• Ictiobius sp. added as species
• Acantopsis sp. added as species

Changes resulting from publication Armbruster (2004)

Citation is: Armbruster, JW. 2004. Phylogenetic relationships of the suckermouth armoured catfishes (Loricariidae) with emphasis on the Hypostominae and the Ancistrinae. Zoological Journal of the Linnean Society 141:1-80.

• Megalancistrus gigas added as synonym for Megalancistrus parananus
• Hemiancistrus panamensis added as synonym for Hemiancistrus aspidolepis
• Hemiancistrus sp. added as species
• Hemiancistrus sp. Brazil added as species
• Peckoltia sp. 1 added as species
• Peckoltia sp. 2 added as species
• Peckoltia sp. big spot added as species
• Pseudancistrus sp. gold spot added as species
• Hypostomus plecostomus 2 added as species
• Hypostomus sp. round snout 1 added as species
• Hypostomus sp. round snout 2 added as species
• Hemipsilichthys sp. added as species
• Hemipsilichthys? added as species
• Kronichthys sp. 1 added as species
• Kronichthys sp. 2 added as species

Changes resulting from publication Schaefer (1990)

Citation is: Schaefer, SA. 1990. Anatomy and relationships of the scoloplacid catfishes. Proceedings of the Academy of Natural Sciences of Philadelphia 142:167-210.

• Arius sp. added as species
• Henonemus sp. added as species
• Stegophilus sp. added as species
• Trichomycterus sp. added as species
• Tridentopsis sp. added as species
• Astroblepus sp. added as species
• Cochliodon sp. added as species
• Neoplecostomus sp. added as species
• Perugia xanthus added as synonym of Megalonema xanthum
• Corydoras barbatus added as synonym of Scieromystax barbatus
• Isorineloricaria spinosissima added as species

Changes resulting from publication Winterbottom (1980)

Citation is: Winterbottom, R. 1980. Systematics, osteology and phylogenetic relationships of fishes of the Ostariophysan subfamily Anostominae (Characoidei, Anostomidae). Royal Ontario Museum Life Science Contributions 123:112.

• Gnathodolus sp. added as species

Changes resulting from publication Castro and Vari (2004)

Citation is: Castro, RMC. and Vari, RP. 2004. Detritivores of the South American fish family Prochilodontidae (Ostariophysi: Characiformes): a phylogenetic and revisionary study. Smithsonian Contributions to Zoology 622:1-190.

• Anostomus sp. added as species

Changes resulting from publication Toledo-Piza (2000)

• Cynodon sp. (Toledo-Piza 2000) added as species
• Hydrolycus sp. (Toledo-Piza 2000) added as species
• Agoniates sp. (Toledo-Piza 2000) added as species
• Gnathocharax sp. (Toledo-Piza 2000) added as species
• Heterocharax sp. (Toledo-Piza 2000) added as species
• Mylossoma sp. (Toledo-Piza 2000) added as species
• Roeboides sp. (Toledo-Piza 2000) added as species
• Serrasalmus sp. (Toledo-Piza 2000) added as species
• Triportheus sp. (Toledo-Piza 2000) added as species
• Hoplias cf. malabaricus (Toledo-Piza 2000) added as species
• Hemiodus sp. (Toledo-Piza 2000) added as species
• Hydrocynus sp. (Toledo-Piza 2000) added as species
• Pyrrhulina sp. (Toledo-Piza 2000) added as species

Changes resulting from publication Toledo-Piza (2007)

Citation is: Toledo-Piza, M. 2007. Phylogenetic relationships among Acestrorhynchus species (Ostariophysi: Characiformes: Acestrorhynchidae). Zoological Journal of the Linnean Society 151:691-757.

• Bryconops sp. (Toledo-Piza 2007) added as species

Changes resulting from publication Lucena and Menezes (1998)

Citation is: Lucena, C. A. S., and N. A. Menezes. 1998. A phylogenetic analysis of Roestes Günther and Gilbertolus Eigenmann with a hypothesis on the relationships of the Cynodontidae and Acestrorhynchidae (Teleostei, Ostariophysi, Characiformes) in L. R. Malabarba, R. E. Reis, R. P. Vari, Z. M. S. Lucena, and C. A. S. Lucena, eds. Phylogeny and Classification of Neotropical Fishes. EDIPUCRS, Porto Alegre, Brazil.

• Brycon sp. (Lucena and Menezes 1989) added as species
• Roeboides sp. (Lucena and Menezes 1989) added as species
• Roestes sp. (Lucena and Menezes 1989) added as species

Changes resulting from publication Weitzman and Fink (1983)

Citation is: Weitzman, S. H., and W. L. Fink. 1983. Relationships of the neon tetras, a group of South American freshwater fishes (Teleostei, Characidae), with comments on the phylogeny of New World Characiforms. Bulletin of the Museum of Comparative Zoology, Harvard University 150:339-395.

• Aphyodite sp. (Weitzman and Fink 1983) added as species
• Characidium sp. (Weitzman and Fink 1983) added as species
• Hemigrammus sp. (Weitzman and Fink 1983) added as species
• Hyphessobrycon sp. (Weitzman and Fink 1983) added as species
• Leptobrycon sp. (Weitzman and Fink 1983) added as species
• Microschemobrycon sp. (Weitzman and Fink 1983) added as species
• Oxybrycon sp. (Weitzman and Fink 1983) added as species
• Pristella sp. (Weitzman and Fink 1983) added as species
• Bryconamericus meridae added as synonym of Knodus meridae'
• Cheirodon piaba added as synonym of Serrapinnus piaba
• Neolebias olbrechtsi added as synonym of Congocharax olbrechtsi

Changes resulting from publication Vari (1983)

Citation is: Vari, R. P. 1983. Phylogenetic relationships of the families Curimatidae, Prochilodontidae, Anostomidae, and Chilodontidae (Pisces: Characiformes). Smithsonian Contributions to Zoology 378:60.

• Ichthyoelephas sp. (Vari 1983) added as species
• Prochilodus sp. (Vari 1983) added as species
• Anostomus sp. (Vari 1983) added as species
• Hemiodus sp. (Vari 1983) added as species
• Citharinus sp. (Vari 1983) added as species

Changes resulting from publication Vari (1989a)

Citation is: Vari, R. P. 1989. A phylogenetic study of the Neotropical characiform family Curimatidae (Pisces: Ostariophysi). Smithsonian Contributions to Zoology 471:iv+1-71.

• Anostomus sp. (Vari 1989a) added as species
• Laemolyta sp. (Vari 1989a) added as species
• Agoniates sp. (Vari 1989a) added as species
• Hollandichthys sp. (Vari 1989a) added as species
• Hydrocynus sp. (Vari 1989a) added as species
• Mesoborus sp. (Vari 1989a) added as species
• Hemiodus sp. (Vari 1989a) added as species
• Ichthyoelephas sp. (Vari 1989a) added as species

Changes resulting from publication Weitzman and Fink (1985)

Citation is: Weitzman, S. H., and S. V. Fink. 1985. Xenurobryconin phylogeny and putative phermone pumps in glandulocaudine fishes (Teleostei: Characidae). Smithsonian Contributions to Zoology ?:1-121.

• Argopleura sp. (Weitzman and Fink 1985) added as species
• Gephyrocharax sp. (Weitzman and Fink 1985) added as species
• Tyttocharax sp. (Weitzman and Fink 1985) added as species

Changes resulting from unpublished thesis of Siebert

Citation is: Siebert, Darrell J. Interrelationships among families of the order Cypriniformes (Teleostei).Unpubl. PhD Thesis, City University of New York. 1987.

• Gyrinocheilus sp. (Siebert 1987) added as species
• Ellopostoma sp. (Siebert 1987) added as species
• Carpiodes sp. (Siebert 1987) added as species
• Ictiobus sp. (Siebert 1987) added as species
• Aopidoparia sp. (Siebert 1987) added as species
• Acheilognathus intermedius added as species
• Pantosteus clarki added as species
• Hybopsis micropogon added as species
• Hybopsis bigutattus added as species
• Barbus matsudai added as species
• Barbus conchonius added as synonym of Puntius conchonius
• Acanthopsis choirorhynchos added as synonym of Acantopsis choirorhynchos
• Acanthophthalmus kuhli added as synonym of Pangio kuhlii
• Chela anomalurus added as synonym of Oxygaster pointoni
• Chondrostoma ohridanus added as synonym of Chondrostoma nasus
• Cirrhinus chinensis added as synonym of Cirrhinus molitorella
• Cyprinion irregularis added as synonym of Cprinion watsoni
• Hemibarbus barbus added as synonym of Hemibarbus labeo
• Epalzeorhynchus kalopterus added as a synonym of Epalzeorhynchos kalopterus
• Labeo erythrura added as a synonym of Epalzeorhynchos frenatum
• Rhynchocypris variegata added as a synonym of Phoxinus oxycephalus
• Varicorhinus tamusuiensis added as a synonym of Varicorhinus barbatulus
• Scaphesthes tamusuiensis added as a synonym of Varicorhinus barbatulus
• Lepidocephalus hasselti added as a synonym of Lepidocephalichthys hasselti
• Leptobotia fasciata added as a synonym of Parabotia fasciata
• Morulius chrysophekadion added as a synonym of Labeo chrysophekadion
• Nemachelus posteroventralis added as a synonym of Barbatula toni
• Noemacheilus selangoricus added as a synonym of Nemacheilus selangoricus
• Hesperoleucas symmetricus added as synonym of Hesperoleucus symmetricus'
• Noemacheilus pulcher added as a synonym of Micronemacheilus pulcher
• Nemacheilus pulcher added as a synonym of Micronemacheilus pulcher

Changes resulting from publication Roberts (1969)

Citation is: Roberts, T. R. 1969. Osteology and relationships of characoid fishes, particularly the genera Hepsetus, Salminus, Hoplias, Ctenolucius and Acestrorhynchus. Proceedings of the California Academy of Sciences 36:391-500.

• Anostomus sp. (Roberts 1969) added as species
• Chilodus sp. (Roberts 1969) added as species
• Hoplias sp. (Roberts 1969) added as species
• Hydrocynus sp. (Roberts 1969) added as species
• Mylossoma sp. (Roberts 1969) added as species
• Serrasalmus sp. (Roberts 1969) added as species

Update from August 2008 Catalog of Fishes

TTO was updated from the Catalog of Fishes release of August 29, 2008 with the TTOupdate tool. In addition to CoF changes, the TTOupdate tool captures intermediate synonyms from the CoF status text for each species name.